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This online database is copyrighted by Species 2000 on behalf of the Catalogue of Life partners.

Use of the content (such as the classification, synonymic species checklist, and scientific names) for publications and databases by individuals and organizations for not-for-profit usage is encouraged, on condition that full and precise credit is given at three levels on all occasions that records are shown. The three-part credit includes the complete work, the contributing database of the record, and the expert who provides taxonomic scrutiny of the individual record. For example, these might be as follows:

1) Species 2000 & ITIS Catalogue of Life: 2013 Annual Checklist,
2) The full or short name of the contributing database, such as: Coreoidea Species File or CoreoideaSF,
3) When provided, the latest taxonomic scrutiny field (specialist name & date), such as Lohrmann V. & Kroupa A.S., 05-Oct-2012 (from HymIS Rhopalosomatidae) or Barber-James H., Sartori M., Gattolliat J.L. & Webb J., Feb-2013 (from FADA Ephemeroptera).

This three-part credit can in many cases be kept brief by using a standard form and one or more logos as in the following example: "Species recognized by J. van Tol, Odonata database, ver. Dec 2011 in CoL_logo". You may copy the Catalogue of Life logo from this example, on the DVD and web versions, for use in giving credit.

If you wish to use the content on a public portal or webpage you are required to notify the Species 2000 Secretariat, both to request written permission and to assist with a check that the correct credits are given.

Commercial Use

Use of this compilation or any of the databases contained within it by commercial organisations requires a written agreement from Species 2000 and ITIS. This applied when it is used commercially for external services, products or publications and when it is used to provide internal services within an organisation.


The Species 2000 & ITIS Catalogue of Life editorial board cannot guarantee the accuracy or completeness of the information in this edition. Be aware that the Catalogue of Life is still incomplete and undoubtedly contains errors. Neither Species 2000 & ITIS nor any contributing database can be made liable for any direct or indirect damage arising out of the use of Catalogue of Life.

Copyright for booklet photographs remains with the contributors.

ISSN 1473-009X